The pair starred together in the hit 1989 film “Say Anything.” Skye writes - almost offhandedly - that years after working together on the project, the two slept together, following her divorce in 2000 from Beastie Boys member Adam “Ad-Rock” Horovitz.

"Was I in love with Johnny?" The actors behind beloved 1989 teen movie Say Anything and characters Diane Court and Lloyd Dobler shared a romance off-screen, too, but it happened years later, actress Ione Skye writes in her new memoir.

The actress talks to PEOPLE about why she decided to tell her life story — including having an abortion with Anthony Kiedis and why her marriage to Beastie Boy Adam Horowitz fell apart — in her memoir 'Say Everything.

The two main actors in the beloved 1989 teen romcom Say Anything, Ione Skye and John Cusack, reportedly slept with each other years after they made the movie. Skye and Cusack played polar opposites who fall in love with each other in Cameron Crowe ’s 1989 film, which has gone on to become a cult classic.

Elsewhere in the memoir, Skye, who began acting at age 16, dished on her short-lived fling with fellow co-star, Matthew Perry, with whom she starred in 1988's A Night in the Life of Jimmy Reardon, alongside River Phoenix. While she originally had a crush on Phoenix, it was her and Perry who would end up getting together years later.

Actress Ione Skye may be best known for the 1989 film Say Anything, co-starring with John Cusack in a sweet love story. Now she’s lifting the veil behind that film, her tryst with her co-star, and several of her other relationships in her new memoir, Say Everything, out Tuesday, March 4.

Say Anything' star Ione Skye spills all on her wild love life with Anthony Kiedis, John Cusack In the domestic diva's taste we trust: From clothing to cookware to $12 priming lotion, these are her affordable faves.